Journal Articles
*Graduate Student, **Postdoc or Research Associate
[1] P. B. Nagelson**, R. A. York, J. J. Kral, C. Phelan**, C. Quirmbach, and S. M. Bisbing, “Treatments mediate wildfire impacts to old-growth giant sequoia,” Forest Ecology and Management, vol. 614, p. 123811, Aug. 2026, doi: 10.1016/j.foreco.2026.123811.
[2] J. Weeks, P. B. Nagelson**, S. Bisbing, and H. Safford, “Burn to Save, or Save to Burn? Management May Be Key to Conservation of an Iconic Old-Growth Stand in California, USA,” Fire, vol. 8, no. 2, p. 70, Feb. 2025, doi: 10.3390/fire8020070.
[3] P. B. Nagelson**, R. A. York, K. T. Shoemaker, D. E. Foster, S. L. Stephens, and S. M. Bisbing, “Repeated fuel treatments fall short of fire‐adapted regeneration objectives in a Sierra Nevada mixed conifer forest, USA,” Ecological Applications, vol. 35, no. 1, p. e3075, Jan. 2025, doi: 10.1002/eap.3075.
[4] D. L. Hankins et al., “Indigenous stewardship rights and opportunities to recenter Indigenous fire,” fire ecol, vol. 21, no. 1, pp. 74, s42408-025-00393–0, Nov. 2025, doi: 10.1186/s42408-025-00393-0.
[5] L. E. Hankin*, F. H. Barrios-Masias, A. K. Urza**, and S. M. Bisbing, “Lethal combination for seedlings: extreme heat drives mortality of drought-exposed high-elevation pine seedlings,” Annals of Botany, vol. 135, no. 1–2, pp. 293–304, Feb. 2025, doi: 10.1093/aob/mcae064.
[6] L. M. Galland, T. M. Faske, C. Osuna‐Mascaró, S. M. Bisbing, and T. L. Parchman, “Geography and Environment Shape Spatial Genetic Variation and Predict Climate Maladaptation Across Isolated and Disjunct Populations of Pinus muricata,” Molecular Ecology, vol. 34, no. 23, p. e17638, Dec. 2025, doi: 10.1111/mec.17638.
[7] S. M. Bisbing, P. B. Nagelson**, L. E. Cox, and C. N. Phelan**, “Post-mortem silviculture meets fuel reduction objective in drought-impacted southern Sierra Nevada forests,” Forest Ecology and Management, vol. 595, p. 123035, Nov. 2025, doi: 10.1016/j.foreco.2025.123035.
[8] J. Ren, E. J. Hanan, P. D’Odorico, C. Tague, J. P. Schimel, and P. M. Homyak, “Dryland Watersheds in Flux: How Nitrogen Deposition and Changing Precipitation Regimes Shape Nitrogen Export,” Earth’s Future, vol. 12, no. 4, p. e2023EF004120, Apr. 2024, doi: 10.1029/2023EF004120.
[9] M. P. North et al., “Strategic fire zones are essential to wildfire risk reduction in the Western United States,” fire ecol, vol. 20, no. 1, pp. 50, s42408-024-00282-y, May 2024, doi: 10.1186/s42408-024-00282-y.
[10] L. E. Hankin*, E. A. Leger, and S. M. Bisbing, “Reforestation of high elevation pines: Direct seeding success depends on seed source and sowing environment,” Ecological Applications, vol. 33, no. 6, p. e2897, Sep. 2023, doi: 10.1002/eap.2897.
[11] E. G. Brodie, E. E. Knapp, A. M. Latimer, H. D. Safford, M. Vossmer, and S. M. Bisbing, “The century-long shadow of fire exclusion: Historical data reveal early and lasting effects of fire regime change on contemporary forest composition,” Forest Ecology and Management, vol. 539, p. 121011, Jul. 2023, doi: 10.1016/j.foreco.2023.121011.
[12] S. M. Bisbing, A. K. Urza**, R. A. York, L. E. Hankin*, and T. R. Putz*, “Persistent, viable seedbank buffers serotinous bishop pine over a broad fire return interval,” fire ecol, vol. 19, no. 1, p. 35, May 2023, doi: 10.1186/s42408-023-00194-3.
[13] S. M. Bisbing, B. J. Buma, B. V. Naald, and A. L. Bidlack, “Single-tree salvage logging as a response to Alaska yellow-cedar climate-induced mortality maintains ecological integrity with limited economic returns,” Forest Ecology and Management, vol. 503, p. 119815, Jan. 2022, doi: 10.1016/j.foreco.2021.119815.
[14] M. C. Agne, J. B. Fontaine, N. J. Enright, S. M. Bisbing, and B. J. Harvey, “Demographic processes underpinning post-fire resilience in California closed-cone pine forests: the importance of fire interval, stand structure, and climate,” Plant Ecol, vol. 223, no. 7, pp. 751–767, Jul. 2022, doi: 10.1007/s11258-022-01228-7.
[15] T. R. Putz*, A. K. Urza**, L. E. Hankin*, and S. M. Bisbing, “Seasonal water availability drives trait variation in isolated Basin and Range Pinus ponderosa,” Forest Ecology and Management, vol. 488, p. 119022, May 2021, doi: 10.1016/j.foreco.2021.119022.
[16] K. M. Nelson**, S. Bisbing, D. L. Grossenbacher, M. Ritter, and J. M. Yost, “Testing an invasion mechanism for Eucalyptus globulus : Is there evidence of allelopathy?,” American J of Botany, vol. 108, no. 4, pp. 607–615, Apr. 2021, doi: 10.1002/ajb2.1635.
[17] K. N. Nelson**, E. O’Dean, E. E. Knapp, A. J. Parker, and S. M. Bisbing, “Persistent yet vulnerable: resurvey of an Abies ecotone reveals few differences but vulnerability to climate change,” Ecology, vol. 102, no. 12, p. e03525, Dec. 2021, doi: 10.1002/ecy.3525.
[18] L. E. Hankin* and S. M. Bisbing, “Let it snow? Spring snowpack and microsite characterize the regeneration niche of high‐elevation pines,” Journal of Biogeography, vol. 48, no. 8, pp. 2068–2084, Aug. 2021, doi: 10.1111/jbi.14136.
[19] S. M. Bisbing, A. K. Urza**, B. J. Buma, D. J. Cooper, M. Matocq, and A. L. Angert, “Can long‐lived species keep pace with climate change? Evidence of local persistence potential in a widespread conifer,” Diversity and Distributions, vol. 27, no. 2, pp. 296–312, Feb. 2021, doi: 10.1111/ddi.13191.
[20] A. L. Bidlack et al., “Climate-Mediated Changes to Linked Terrestrial and Marine Ecosystems across the Northeast Pacific Coastal Temperate Rainforest Margin,” BioScience, vol. 71, no. 6, pp. 581–595, Jun. 2021, doi: 10.1093/biosci/biaa171.
[21] B. Buma et al., “Emergent freeze and fire disturbance dynamics in temperate rainforests,” Austral Ecology, vol. 44, no. 5, pp. 812–826, Aug. 2019, doi: 10.1111/aec.12751.
[22] B. Buma, S. M. Bisbing, G. Wiles, and A. L. Bidlack, “100 yr of primary succession highlights stochasticity and competition driving community establishment and stability,” Ecology, vol. 100, no. 12, p. e02885, Dec. 2019, doi: 10.1002/ecy.2885.
[23] S. M. Bisbing, B. J. Buma, L. E. Oakes, J. Krapek, and A. L. Bidlack, “From canopy to seed: Loss of snow drives directional changes in forest composition,” Ecology and Evolution, vol. 9, no. 14, pp. 8157–8174, Jul. 2019, doi: 10.1002/ece3.5383.
[24] S. M. Bisbing and D. V. D’Amore, “Nitrogen dynamics vary across hydrologic gradients and by forest community composition in the perhumid coastal temperate rainforest of southeast Alaska,” Can. J. For. Res., vol. 48, no. 2, pp. 180–191, Feb. 2018, doi: 10.1139/cjfr-2017-0178.
[25] M. Petrášová-Šibíková et al., “Effect of the Gabčíkovo Waterworks (Slovakia) on riparian floodplain forest ecosystems in the Danube inland delta: vegetation dynamics and trends,” Biologia, vol. 72, no. 7, pp. 722–734, Jul. 2017, doi: 10.1515/biolog-2017-0082.
[26] M. Petrášová-Šibíková et al., “Effect of the Gabčíkovo Waterworks (Slovakia) on riparian floodplain forest ecosystems in the Danube inland delta: vegetation dynamics and trends,” Biologia, vol. 72, no. 7, pp. 722–734, Jul. 2017, doi: 10.1515/biolog-2017-0082.
[27] A. Bidlack et al., “Alternative interpretation and scale-based context for ‘No evidence of recent (1995–2013) decrease of yellow-cedar in Alaska’ (Barrett and Pattison 2017),” Can. J. For. Res., vol. 47, no. 8, pp. 1145–1151, Aug. 2017, doi: 10.1139/cjfr-2017-0070.
[28] R. L. Mulvey and S. M. Bisbing, “Complex Interactions Among Agents Affect Shore Pine Health in Southeast Alaska,” Northwest Science, vol. 90, no. 2, pp. 176–194, May 2016, doi: 10.3955/046.090.0209.
[29] S. M. Bisbing, D. J. Cooper, D. V. D’Amore, and K. N. Marshall, “Determinants of conifer distributions across peatland to forest gradients in the coastal temperate rainforest of southeast Alaska,” Ecohydrology, vol. 9, no. 2, pp. 354–367, Mar. 2016, doi: 10.1002/eco.1640.
[30] M. Budzakova, I. Hodálová, P. Mereďa, L. Somlyay, and S. M. Bisbing, “Karyological, morphological, and ecological differentiation of Sesleria caerula and Sesleria tatrae (Poaceae) in the Western Carpathians and adjacent regions,” Preslia, vol. 86, pp. 245–277, 2014.
[31] H. Bothwell et al., “Identifying genetic signatures of selection in a non-model species, alpine gentian (Gentiana nivalis L.), using a landscape genetic approach,” Conserv Genet, vol. 14, no. 2, pp. 467–481, Apr. 2013, doi: 10.1007/s10592-012-0411-5.
[32] S. M. Bisbing, P. B. Alaback, and T. H. DeLuca, “Carbon storage in old-growth and second growth fire-dependent western larch (Larix occidentalis Nutt.) forests of the Inland Northwest, USA,” Forest Ecology and Management, vol. 259, no. 5, pp. 1041–1049, Feb. 2010, doi: 10.1016/j.foreco.2009.12.018.
[33] S. M. Bisbing, P. B. Alaback, and T. H. DeLuca, “Carbon storage in old-growth and second growth fire-dependent western larch (Larix occidentalis Nutt.) forests of the Inland Northwest, USA,” Forest Ecology and Management, vol. 259, no. 5, pp. 1041–1049, Feb. 2010, doi: 10.1016/j.foreco.2009.12.018.
[2] J. Weeks, P. B. Nagelson**, S. Bisbing, and H. Safford, “Burn to Save, or Save to Burn? Management May Be Key to Conservation of an Iconic Old-Growth Stand in California, USA,” Fire, vol. 8, no. 2, p. 70, Feb. 2025, doi: 10.3390/fire8020070.
[3] P. B. Nagelson**, R. A. York, K. T. Shoemaker, D. E. Foster, S. L. Stephens, and S. M. Bisbing, “Repeated fuel treatments fall short of fire‐adapted regeneration objectives in a Sierra Nevada mixed conifer forest, USA,” Ecological Applications, vol. 35, no. 1, p. e3075, Jan. 2025, doi: 10.1002/eap.3075.
[4] D. L. Hankins et al., “Indigenous stewardship rights and opportunities to recenter Indigenous fire,” fire ecol, vol. 21, no. 1, pp. 74, s42408-025-00393–0, Nov. 2025, doi: 10.1186/s42408-025-00393-0.
[5] L. E. Hankin*, F. H. Barrios-Masias, A. K. Urza**, and S. M. Bisbing, “Lethal combination for seedlings: extreme heat drives mortality of drought-exposed high-elevation pine seedlings,” Annals of Botany, vol. 135, no. 1–2, pp. 293–304, Feb. 2025, doi: 10.1093/aob/mcae064.
[6] L. M. Galland, T. M. Faske, C. Osuna‐Mascaró, S. M. Bisbing, and T. L. Parchman, “Geography and Environment Shape Spatial Genetic Variation and Predict Climate Maladaptation Across Isolated and Disjunct Populations of Pinus muricata,” Molecular Ecology, vol. 34, no. 23, p. e17638, Dec. 2025, doi: 10.1111/mec.17638.
[7] S. M. Bisbing, P. B. Nagelson**, L. E. Cox, and C. N. Phelan**, “Post-mortem silviculture meets fuel reduction objective in drought-impacted southern Sierra Nevada forests,” Forest Ecology and Management, vol. 595, p. 123035, Nov. 2025, doi: 10.1016/j.foreco.2025.123035.
[8] J. Ren, E. J. Hanan, P. D’Odorico, C. Tague, J. P. Schimel, and P. M. Homyak, “Dryland Watersheds in Flux: How Nitrogen Deposition and Changing Precipitation Regimes Shape Nitrogen Export,” Earth’s Future, vol. 12, no. 4, p. e2023EF004120, Apr. 2024, doi: 10.1029/2023EF004120.
[9] M. P. North et al., “Strategic fire zones are essential to wildfire risk reduction in the Western United States,” fire ecol, vol. 20, no. 1, pp. 50, s42408-024-00282-y, May 2024, doi: 10.1186/s42408-024-00282-y.
[10] L. E. Hankin*, E. A. Leger, and S. M. Bisbing, “Reforestation of high elevation pines: Direct seeding success depends on seed source and sowing environment,” Ecological Applications, vol. 33, no. 6, p. e2897, Sep. 2023, doi: 10.1002/eap.2897.
[11] E. G. Brodie, E. E. Knapp, A. M. Latimer, H. D. Safford, M. Vossmer, and S. M. Bisbing, “The century-long shadow of fire exclusion: Historical data reveal early and lasting effects of fire regime change on contemporary forest composition,” Forest Ecology and Management, vol. 539, p. 121011, Jul. 2023, doi: 10.1016/j.foreco.2023.121011.
[12] S. M. Bisbing, A. K. Urza**, R. A. York, L. E. Hankin*, and T. R. Putz*, “Persistent, viable seedbank buffers serotinous bishop pine over a broad fire return interval,” fire ecol, vol. 19, no. 1, p. 35, May 2023, doi: 10.1186/s42408-023-00194-3.
[13] S. M. Bisbing, B. J. Buma, B. V. Naald, and A. L. Bidlack, “Single-tree salvage logging as a response to Alaska yellow-cedar climate-induced mortality maintains ecological integrity with limited economic returns,” Forest Ecology and Management, vol. 503, p. 119815, Jan. 2022, doi: 10.1016/j.foreco.2021.119815.
[14] M. C. Agne, J. B. Fontaine, N. J. Enright, S. M. Bisbing, and B. J. Harvey, “Demographic processes underpinning post-fire resilience in California closed-cone pine forests: the importance of fire interval, stand structure, and climate,” Plant Ecol, vol. 223, no. 7, pp. 751–767, Jul. 2022, doi: 10.1007/s11258-022-01228-7.
[15] T. R. Putz*, A. K. Urza**, L. E. Hankin*, and S. M. Bisbing, “Seasonal water availability drives trait variation in isolated Basin and Range Pinus ponderosa,” Forest Ecology and Management, vol. 488, p. 119022, May 2021, doi: 10.1016/j.foreco.2021.119022.
[16] K. M. Nelson**, S. Bisbing, D. L. Grossenbacher, M. Ritter, and J. M. Yost, “Testing an invasion mechanism for Eucalyptus globulus : Is there evidence of allelopathy?,” American J of Botany, vol. 108, no. 4, pp. 607–615, Apr. 2021, doi: 10.1002/ajb2.1635.
[17] K. N. Nelson**, E. O’Dean, E. E. Knapp, A. J. Parker, and S. M. Bisbing, “Persistent yet vulnerable: resurvey of an Abies ecotone reveals few differences but vulnerability to climate change,” Ecology, vol. 102, no. 12, p. e03525, Dec. 2021, doi: 10.1002/ecy.3525.
[18] L. E. Hankin* and S. M. Bisbing, “Let it snow? Spring snowpack and microsite characterize the regeneration niche of high‐elevation pines,” Journal of Biogeography, vol. 48, no. 8, pp. 2068–2084, Aug. 2021, doi: 10.1111/jbi.14136.
[19] S. M. Bisbing, A. K. Urza**, B. J. Buma, D. J. Cooper, M. Matocq, and A. L. Angert, “Can long‐lived species keep pace with climate change? Evidence of local persistence potential in a widespread conifer,” Diversity and Distributions, vol. 27, no. 2, pp. 296–312, Feb. 2021, doi: 10.1111/ddi.13191.
[20] A. L. Bidlack et al., “Climate-Mediated Changes to Linked Terrestrial and Marine Ecosystems across the Northeast Pacific Coastal Temperate Rainforest Margin,” BioScience, vol. 71, no. 6, pp. 581–595, Jun. 2021, doi: 10.1093/biosci/biaa171.
[21] B. Buma et al., “Emergent freeze and fire disturbance dynamics in temperate rainforests,” Austral Ecology, vol. 44, no. 5, pp. 812–826, Aug. 2019, doi: 10.1111/aec.12751.
[22] B. Buma, S. M. Bisbing, G. Wiles, and A. L. Bidlack, “100 yr of primary succession highlights stochasticity and competition driving community establishment and stability,” Ecology, vol. 100, no. 12, p. e02885, Dec. 2019, doi: 10.1002/ecy.2885.
[23] S. M. Bisbing, B. J. Buma, L. E. Oakes, J. Krapek, and A. L. Bidlack, “From canopy to seed: Loss of snow drives directional changes in forest composition,” Ecology and Evolution, vol. 9, no. 14, pp. 8157–8174, Jul. 2019, doi: 10.1002/ece3.5383.
[24] S. M. Bisbing and D. V. D’Amore, “Nitrogen dynamics vary across hydrologic gradients and by forest community composition in the perhumid coastal temperate rainforest of southeast Alaska,” Can. J. For. Res., vol. 48, no. 2, pp. 180–191, Feb. 2018, doi: 10.1139/cjfr-2017-0178.
[25] M. Petrášová-Šibíková et al., “Effect of the Gabčíkovo Waterworks (Slovakia) on riparian floodplain forest ecosystems in the Danube inland delta: vegetation dynamics and trends,” Biologia, vol. 72, no. 7, pp. 722–734, Jul. 2017, doi: 10.1515/biolog-2017-0082.
[26] M. Petrášová-Šibíková et al., “Effect of the Gabčíkovo Waterworks (Slovakia) on riparian floodplain forest ecosystems in the Danube inland delta: vegetation dynamics and trends,” Biologia, vol. 72, no. 7, pp. 722–734, Jul. 2017, doi: 10.1515/biolog-2017-0082.
[27] A. Bidlack et al., “Alternative interpretation and scale-based context for ‘No evidence of recent (1995–2013) decrease of yellow-cedar in Alaska’ (Barrett and Pattison 2017),” Can. J. For. Res., vol. 47, no. 8, pp. 1145–1151, Aug. 2017, doi: 10.1139/cjfr-2017-0070.
[28] R. L. Mulvey and S. M. Bisbing, “Complex Interactions Among Agents Affect Shore Pine Health in Southeast Alaska,” Northwest Science, vol. 90, no. 2, pp. 176–194, May 2016, doi: 10.3955/046.090.0209.
[29] S. M. Bisbing, D. J. Cooper, D. V. D’Amore, and K. N. Marshall, “Determinants of conifer distributions across peatland to forest gradients in the coastal temperate rainforest of southeast Alaska,” Ecohydrology, vol. 9, no. 2, pp. 354–367, Mar. 2016, doi: 10.1002/eco.1640.
[30] M. Budzakova, I. Hodálová, P. Mereďa, L. Somlyay, and S. M. Bisbing, “Karyological, morphological, and ecological differentiation of Sesleria caerula and Sesleria tatrae (Poaceae) in the Western Carpathians and adjacent regions,” Preslia, vol. 86, pp. 245–277, 2014.
[31] H. Bothwell et al., “Identifying genetic signatures of selection in a non-model species, alpine gentian (Gentiana nivalis L.), using a landscape genetic approach,” Conserv Genet, vol. 14, no. 2, pp. 467–481, Apr. 2013, doi: 10.1007/s10592-012-0411-5.
[32] S. M. Bisbing, P. B. Alaback, and T. H. DeLuca, “Carbon storage in old-growth and second growth fire-dependent western larch (Larix occidentalis Nutt.) forests of the Inland Northwest, USA,” Forest Ecology and Management, vol. 259, no. 5, pp. 1041–1049, Feb. 2010, doi: 10.1016/j.foreco.2009.12.018.
[33] S. M. Bisbing, P. B. Alaback, and T. H. DeLuca, “Carbon storage in old-growth and second growth fire-dependent western larch (Larix occidentalis Nutt.) forests of the Inland Northwest, USA,” Forest Ecology and Management, vol. 259, no. 5, pp. 1041–1049, Feb. 2010, doi: 10.1016/j.foreco.2009.12.018.
Proceedings
Brousil, M.* and S. Bisbing 2017, "Influence of fire history on coast redwood regeneration and stand structure." In Standiford and Valachovic. Proceedings of the Coast Redwood Science Symposium. Gen. Tech. Rep. PSW-GTR-258.